Hi! That is indeed true, because the Zynthian software has to perform an internal resampling, to adapt the digital data of the waveforms before they hit the converters of the audio board, at the set sample rate.
We should also remember that the recent slew of low-level software emulators of digital synthesis architectures - made possible by computational power increased by orders of magnitude, with respect to the past - is based on exact replications of the DSP workings with their running DSP code.
This also means that nothing of what possibly surrounds or follows the signal chain of the digital sound, whose last stage are the DACs, is mirrored in the emulatorās audio output.
For example, the JV-880 was cleverly engineered with a simple and inexpensive post-DAC discrete circuitry stage, made of an op-amp and an elementary but effective fixed-freq 12 dB/Oct LP filter, set at about 15500 Hz, that is exactly at the upper threshold of the available bandwidth of 16 Khz at 32 Khz of sample rate.
This cheap hardware provision imparted analog softness, air and delicacy, to the upper range of the digital spectrum, thus masking the unavoidable digital artifacts of quantisation, and possibly any residual audio bleed from the anti-aliasing filterās carrier frequency.
I guess that in later implementations of digital synthesis, like the mid-90s and ensuing VA engines, the impact of any discrete circuitry possibly present after the DAC stage was less impacting on the sonic quality, due to higher DSP power and better converters. Hence, the exceptional results - mostly identical to the original hardware - of the various DSP emulators of a later generation of VA synths (among which the ones from T.U.S., included in the Oram installation).
Anyway, for anyone like me keen of the 1990s catalogue of the JV/JD/XV manufacturer, this is the Don Solarisā widely acknowledged Internet Bible, for tech specs of these venerable and still beloved instruments: